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Third Age of the Eocene Epoch
The Bartonian is, in the International Commission on Stratigraphy's (ICS) geologic time scale, a stage or age in the middle of the Eocene Epoch or Series
Bartonian
Family of aquatic flightless birds
dated to 42 mya. An unnamed fossil from Argentina proves that, by the Bartonian (Middle Eocene), some 39–38 mya, primitive penguins had spread to South
Penguin
Extinct genus of reptiles
Cuvieribaena is an extinct genus of blanid that lived during the Bartonian stage of the Eocene epoch. Cuvieribaena carlgansi is known from France. Čerňanský
Cuvieribaena
Second age of the Eocene Epoch
Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene Subepoch
Lutetian
Extinct genus of carnivorous ungulate from Eocene epoch
Mongolia, each dated to the Irdinmanhan Asian land mammal age (Lutetian–Bartonian stages, 48–38 million years ago). Andrewsarchus has historically been
Andrewsarchus
Extinct genus of sparassodont
genus of sparassodont that lived in South America during the Lutetian and Bartonian stages of the Eocene epoch. Silvenator brasiliensis, a taxon known from
Nemolestes
Third epoch of the Paleogene Period
biosphere, being part of a broader trend of global cooling lasting from the Bartonian to the Rupelian. The transition is marked by the Oi1 event, an oxygen
Oligocene
Extinct genus of mammals
Arenagale is an extinct genus of tenrecid that lived during the Bartonian stage of the Eocene epoch. Arenagale calcareus is known from the Eocliff Limestone
Arenagale
Extinct genus of suliform bird
is an extinct monotypic genus of suliform bird that lived during the Bartonian stage of the Eocene epoch. The generic name of Mangystania refers to the
Mangystania
Second epoch of the Paleogene Period
constitutes the lower, the Priabonian Stage the upper; and the Lutetian and Bartonian stages are united as the middle Eocene.[citation needed] The Western North
Eocene
Extinct genus of borhyaenid sparassodont
genus of borhyaenid sparassodont that lived in South America during the Bartonian stage of the Eocene epoch. The generic name Angelocabrerus honours the
Angelocabrerus
Genus of mammals
Argorheomys is an extinct genus of pipestoneomyid rodent that lived during the Bartonian stage of the Eocene epoch. Argorheomys septendrionalis is known from Duchesnean
Argorheomys
Extinct genus of reptiles
deposited prior to the Bartonian, with a mammalian tooth from the locality suggesting an age between the middle Lutetian and Bartonian, while the layers directly
Dentaneosuchus
Extinct genus of early cetacean
is an extinct genus of an early whale from Peru that lived during the Bartonian age of the middle Eocene. Perucetus is the largest Eocene whale, with
Perucetus
Extinct genus of basilosaurid whales
Tutcetus is an extinct genus of diminutive basilosaurid cetacean from the Bartonian of Egypt. Tutcetus, named after the child pharaoh Tutankhamun, is both
Tutcetus
Extinct genus of primates
Amamria is an extinct genus of primate that lived during the Bartonian stage of the Eocene epoch. Amamria tunisiensis is known from Djebel el Kébar in
Amamria
Extinct genus of fishes
Azerbaijan †Anenchelum paucivertebrale Bannikov & Parin, 1995 - Middle Eocene (Bartonian) of Krasnodar, Russia (Kuma Formation) The species Lepidopus brevicauda
Anenchelum
Extinct genus of whales
best known remains generally suggest that Pachycetus lived during the Bartonian, however, fossil finds have also been recovered from sediments of less
Pachycetus
Village in Hampshire, England
the Barton geological beds. The Barton Beds may give their name to the Bartonian stage of the Eocene Epoch of geological time. The beach is separated from
Barton_on_Sea
American vertebrate palaeontologist
September 2015). Friedman, Matt (ed.). "A New Large Hyainailourine from the Bartonian of Europe and Its Bearings on the Evolution and Ecology of Massive Hyaenodonts
Matthew_Borths
Tukwila Formation Stratigraphic range: Bartonian Type Geological formation Location Region Washington Country United States Type section Named for Tukwila
Tukwila_Formation
Family of ray-finned fishes
Iran. An older, indeterminate Rybapina specimen from the middle Eocene (Bartonian) of the North Caucasus represents the earliest record of the group. Cubariomma
Driftfish
Geologic formation in France
Molasses du Castrais, is a geologic formation of Eocene (Ypresian to Bartonian) age, outcropping in the Montagne-Noire region of France. "Petrified fossil
Sables_du_Castrais_Formation
Fourth and last Age of the Eocene Epoch
the time between 37.71 and 33.9 Ma. The Priabonian is preceded by the Bartonian and is followed by the Rupelian, the lowest stage of the Oligocene. Priabona
Priabonian
Family of ray-finned fishes
Oligocene of Switzerland †Gornylistes Bannikov & Tyler, 2008 - Middle Eocene (Bartonian) of the North Caucasus, Russia †Lobodus Costa, 1866 - Late Eocene to Early
Triggerfish
Extinct genus of whales
genus of basilosaurid early whale that lived during the Late Eocene (Bartonian-Priabonian, 40.4 to 33.9 million years ago.) Specimens have been found
Cynthiacetus
First period of the Cenozoic Era
stages: the Ypresian 56.0 Ma to 47.8 Ma; Lutetian 47.8 Ma to 41.2 Ma; Bartonian 41.2 Ma to 37.71 Ma; and, Priabonian 37.71 Ma to 33.9 Ma. The GSSP for
Paleogene
Extinct genus of sea snails
tokodensis Kovács and Vicián, 2020. Size: 72 mm. Upper Lutetian–Lower Bartonian grey marl (Eocene), Tokod (Hungary). Hungarian Natural History Museum
Harmatia
Extinct genus of galagid primate
monotypic genus of strepsirrhine primate that lived in Afro-Arabia during the Bartonian stage of the Eocene epoch. The generic name Saharagalago references the
Saharagalago
Extinct species of fish
from the Early Eocene of Italy and S. natgeosoc from the Middle Eocene (Bartonian) of North Caucasus, Russia. The species Hemicaranx archaeus, also found
Seriola_sanctaebarbarae
Extinct genus of birds
Lithornis is a genus of extinct paleognathous birds. Although Lithornis was able to fly well, their closest relatives are the extant tinamous (which are
Lithornis
First age of the Paleocene Epoch
Eocene Oligo. MC Maastricht. Danian Selandian Thanetian Ypresian Lutetian Bartonian Priabonian Rupelian Chattian Aquitanian ← PETM ← First Antarctic permanent
Danian
Extinct species of gastropod
family Viviparidae, the river snails. It lived between the Bathonian and Bartonian epochs; although it may have persisted into the Oligocene. Viviparus cariniferus
Viviparus_cariniferus
Extinct genus of mammals
(2): e1. doi:10.18563/pv.40.2.e1. J. J. Hooker. 1986. Mammals from the Bartonian (middle/late Eocene) of the Hampshire Basin, southern England. Bulletin
Propalaeotherium
Extinct genus of mammals
Sperrgale is an extinct genus of tenrecid that lived during the Bartonian stage of the Eocene epoch. Sperrgale minutus is known from the Eocliff Limestone
Sperrgale
Superfamily of mammals
Ursoidea Temporal range: Bartonian - Present 37–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N Life reconstruction of Cephalogale shareri Brown bear (Ursus arctos)
Ursoidea
Extinct genus of scincid lizard
extinct monotypic genus of scincid lizard that lived in France during the Bartonian stage of the Eocene epoch. The generic name Lavergnesaurus is derived
Lavergnesaurus
Zouhri. 2015. New fauna of archaeocete whales (Mammalia, Cetacea) from the Bartonian middle Eocene of southern Morocco. Journal of African Earth Sciences 111:273-286
List of fossiliferous stratigraphic units in Western Sahara
List_of_fossiliferous_stratigraphic_units_in_Western_Sahara
Unit Location Notes Images Eocetus Nom. nov. Valid Fraas Middle Eocene (Bartonian) Giushi Formation Egypt Morocco A protocetid. Replacement name for Mesocetus
1904_in_paleontology
Subfamily of mammals
that lived during the middle Eocene. The best-dated remains stem from Bartonian strata, but some finds suggest that they could have first appeared during
Pachycetinae
Genus of ray-finned fishes
known in the extinct †Eochampsodon Bannikov, 2004 from the Middle Eocene (Bartonian) of the North Caucasus, Russia. The currently recognized species in this
Champsodon
First Age of the Oligocene Epoch
Eocene Oligo. MC Maastricht. Danian Selandian Thanetian Ypresian Lutetian Bartonian Priabonian Rupelian Chattian Aquitanian ← PETM ← First Antarctic permanent
Rupelian
Extinct genus of anthropoid primate
the Bartonian stage of the Eocene epoch. Saharopithecus salemi is known from the Dur At-Talah fossil site of Libya, dating back to the Bartonian stage
Saharopithecus
Extinct genus of mammals
Rupestrohyrax is an extinct genus of titanohyracid that lived during the Bartonian stage of the Eocene epoch. Rupestohyrax palustris is known from the Eoridge
Rupestrohyrax
Sedimentary basin in the South Atlantic Ocean
Pliocene Messinian Tortonian Serravallian Langhian Chattian Rupelian Bartonian Priabonian Lutetian Ypresian Paleocene Maastrichtian Campanian Santonian
Pelotas_Basin
2003. The college also published an annual BarTalk until 1994, when the Bartonian was published for one year. Previously, the Barton Common Room was filled
Barton College (Deakin University)
Barton_College_(Deakin_University)
Extinct genus of turtles
magnifica (Hutchison et al., 2004) Pandaung Formation, Myanmar, Late Eocene (Bartonian) Allaeochelys meylani (Rollot, 2025) Moghra Formation, Egypt, Miocene
Allaeochelys
British teacher and local councillor
particularly during the COVID-19 pandemic in England. "Truly Honoured". Bartonian: Local News and Views from Barton Town Council. March 2022. "A hotelier
Margaret_Elizabeth_Sidell
species is A. heckmani. Hadrianus schucherti Sp nov Jr synonym Hay Eocene Bartonian Jackson Formation US Alabama First described as a Hadrianus species. Moved
1899_in_paleontology
Extinct genus of turtle
extinct genus of cheloniid turtle that lived in Catalonia during the Bartonian. It is a monotypic genus known from a single species, O. decorata. de
Osonachelus
Geological formation in Nigeria
The Ameki Formation is a Lutetian to Bartonian geological formation located in Nigeria. It belongs to the Bende-Ameki Group, which also consists of two
Ameki_Formation
Family of mammals
Canidae (/ˈkæn.ɪ.diː/; from Latin, canis, "dog") is a biological family of caniform carnivorans, constituting a clade. A member of this family is a canid
Canidae
Extinct species of crab
2007 from a coastal exposure of foraminiferal limestone dating to the Bartonian Eocene on the island of Hvar in Croatia. Harpactocarcinus dalmatius has
Harpactocarcinus_dalmatius
Town in North Lincolnshire, England
dark side of life". Yorkshire Post. 8 November 2017. "David Mason MBE". Bartonian: News and Views from Barton Town Council. March 2022. p. 15. "Messo, George"
Barton-upon-Humber
Extinct genus of mammals
Namachloris is an extinct genus of chrysochlorid that lived during the Bartonian stage of the Eocene epoch. Namachloris arenatans is known from the Eocliff
Namachloris
Global warming about 55 million years ago
Neogene Paleo. Eocene Oligo. Danian Selandian Thanetian Ypresian Lutetian Bartonian Priabonian Rupelian Chattian ← PETM ← First Antarctic permanent ice-sheets
Paleocene–Eocene thermal maximum
Paleocene–Eocene_thermal_maximum
Maximum (METM), was a period of very warm climate that occurred during the Bartonian stage, from around 40.5 to 40.0 Ma. It marked a notable reversal of the
Middle Eocene Climatic Optimum
Middle_Eocene_Climatic_Optimum
Genus of mammals
38–34 Ma) of Louisiana, Alabama, and Mississippi, United States, and the Bartonian (43–37 Ma on the New Zealand geologic time scale) to the late Eocene of
Zygorhiza
Genus of mammals
symphysis, a deciduous premolar, and unerupted molars. It was found in Bartonian (40.4 to 37.2 million years ago) layers of the Ameki Formation (6°00′N
Pappocetus
Oxyaenid genus from upper Eocene Epoch
Sarkastodon Temporal range: Middle Eocene (Lutetian to Bartonian) 45.0–39.9 Ma PreꞒ Ꞓ O S D C P T J K Pg N Skull reconstructions of Sarkastodon mongoliensis
Sarkastodon
Extinct genus of birds
Namapsitta is an extinct genus of psittaciform that lived during the Bartonian stage of the Eocene epoch. Namapsitta praeruptorum is known from the Eocliff
Namapsitta
System that relates geologic strata to time
and raptors evolve. Basilosaurus is first fully aquatic whale. 37.71 * Bartonian 41.03 Lutetian 48.07 * Ypresian Greenland separates from Eurasia and Eurasian
Geologic_time_scale
Genus of deep sea hatchetfishes
Iran) The earliest unequivocal fossils of Argyropelecus date to the Bartonian stage of the Eocene. Previous to their discovery, the Eocene fossil record
Argyropelecus
Genus of gastropods
(2010). "New species of Neritidae (Neritimorpha) from the Ypresian and Bartonian of the Paris and Basse-Loire Basins, France". Zootaxa 2606: 55-68. von
Neritodryas
Prehistoric devil ray genus
show teeth identical to B. daviesi coexisting with Eoplinthicus in the Bartonian. A Ypresian species, B. nessovi, known from a singular site in Kazakhstan
Burnhamia_(devil_ray)
Extinct family of mammals
of Paroxyclaenidae in Europe would be Paravulpavoides cooperi from the Bartonian of England with this being one of the smallest members of the family (around
Paroxyclaenidae
Species of mammal (fossil)
is an extinct genus of basilosaurid cetacean from the Middle Eocene (Bartonian stage) Paracas Formation of Peru. It has been noted for its relatively
Supayacetus
Extinct genus of mammals
Namagale is an extinct genus of potamogalid that lived during the Bartonian stage of the Eocene epoch. Namagale grandis is known from the Eocliff Limestone
Namagale
Impact crater in northern Canada
Haughton impact crater is located on Devon Island, Nunavut in far Northern Canada. It is about 23 km (14 mi) in diameter and was formed 31–32 million years
Haughton_impact_crater
Extinct genus of rodents
Pozomys is an extinct genus of caviomorph that lived during the Bartonian stage of the Eocene epoch. Pozomys ucayaliensis is known from the Pozo Formation
Pozomys
Network of subterranean mines under Paris, France
[citation needed] Paris's next important mineral deposit came with the Bartonian age. After a period of land-sea alternation that brought layers of sand
Mines_of_Paris
Genus of mammals
cetacean known from fossils recovered from the Late Eocene sediments (Bartonian-Priabonian stages) of the southeastern United States (most notably Alabama
Pontogeneus
Geologic formation in Morocco
shallow coastal environment. The formation represents the world's richest Bartonian archaic whale assemblage, including protocetids and basilosaurids. Includes
Aridal_Formation
Extinct genus of birds
Scopelortyx Temporal range: Bartonian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum: Chordata Class: Aves Order: Galliformes
Scopelortyx
Extinct family of mammals
doi:10.1016/S0031-0182(00)00252-2. J. J. Hooker. 1986. Mammals from the Bartonian (middle/late Eocene) of the Hampshire Basin, southern England. Bulletin
Xiphodontidae
Geologic formation in Curaçao
geologic formation in Curaçao. It preserves fossils dating back to the Bartonian period. List of fossiliferous stratigraphic units in Curaçao Ceru di Cueba
Ceru_di_Cueba_Formation
Family of fishes
containing the species E. bimaxillaria, known from the mid/late Eocene (Bartonian) of the North Caucasus. The fossil genus Austromola containing one species
Molidae
Geological formation and major fossil site in Antarctica
to just the older Thanetian to Lutetian-aged strata, with the younger Bartonian to Rupelian strata treated as the overlying Submeseta Formation. However
La_Meseta_Formation
Family of single-celled organisms
Cassigerinellidae Temporal range: Bartonian - Serravallian Scientific classification Domain: Eukaryota Clade: Sar Clade: Rhizaria Phylum: Retaria Subphylum:
Cassigerinellidae
Extinct protocetid early whale
is an extinct protocetid early whale known from the early-late Eocene (Bartonian, 40.4 to 37.2 million years ago) Giushi Formation in Gebel Mokattam, (30°00′N
Eocetus
Extinct genus of fishes
contains a single species, E. holosquamatus from the mid-late Eocene (late Bartonian or early Priabonian)-aged Burside Mudstone Formation of the South Island
Eothyrsites
Extinct genus of reptiles
dominance of large reptiles in Europe, at least, had subsided by the Bartonian stage of the Eocene. Increased competition, reduction in prey availability
Barinasuchus
Extinct genus of mammals
Dzavui Temporal range: 41.3–28.1 Ma Bartonian to the top of the Rupelian PreꞒ Ꞓ O S D C P T J K Pg N Scientific classification Kingdom: Animalia Phylum:
Dzavui_(mammal)
Commune in Nouvelle-Aquitaine, France
composed of lower molasse (predominantly clayey facies) (Upper Continental Bartonian). The most recent formation, designated CFp, is one of the undifferentiated
Gageac-et-Rouillac
Prehistoric cetacean genus from the Late Eocene epoch
(2015). "New fauna of archaeocete whales (Mammalia, Cetacea) from the Bartonian middle Eocene of southern Morocco". Journal of African Earth Sciences
Basilosaurus
Family of mammals
Qaisracetus Rodhocetus Takracetus Togocetus Georgiacetinae- Georgiacetines are Bartonian protocetids, considered transitional to the basilosaurids. Their skulls
Protocetidae
Extinct genus of fishes
known: †E. gornylutshensis Bannikov & Tyler, 2008 - mid-late Eocene (Bartonian) of the North Caucasus, Russia (Kuma Formation) †E. pygmaeus (de Zigno
Eotetraodon
Public land-grant university in College Station, Texas, US
Relations. March 17, 1998. Retrieved December 3, 2021. "Front Matter". The Bartonian Legacy. Published by Imperial College Press and Distributed by World Scientific
Texas_A&M_University
Extinct subfamily of mammals
Michael Morlo; Katharina Bastl (2015). "A New Large Hyainailourine from the Bartonian of Europe and Its Bearings on the Evolution and Ecology of Massive Hyaenodonts
Hyainailourinae
Extinct genus of shark
of Austria, Germany & Japan R. fiedleri (Pfeil, 1983) – Late Eocene (Bartonian) of Austria R. goliath (Antunes & Cappetta, 2002) – Late Cretaceous (Campanian)
Rolfodon
Extinct genus of endemic Palaeogene European artiodactyls
University of Montpellier. Hooker, Jerry J. (1986). "Mammals from the Bartonian (middle late Eocene) of the Hampshire Basin, southern England". Bulletin
Dacrytherium
Extinct genus of Devilray
season in which this species was first collected. It is known from the Bartonian-aged Souar-Fortuna formations of Tunisia and possibly the late Eocene
Amamriabatis
villeveyracensis. Lavergnesaurus Gen. et sp. nov Valid Čerňanský et al. Eocene (Bartonian) Phosphorites du Quercy France A skink. The type species is L. lamarcki
2026_in_reptile_paleontology
Extinct genus of fishes
(Dabakhan Formation) P. pshekhaensis Bannikov, 1993 - middle Eocene (Bartonian) of Krasnodar Krai, Russia (Kuma Formation) P. stolyarovi Bratishko &
Palimphyes
Extinct species of shark
(Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths".
Galeocerdo_latidens
Family of small mammals
Tenrecidae Temporal range: Bartonian–Recent PreꞒ Ꞓ O S D C P T J K Pg N Tailless tenrec (Tenrec ecaudatus) Scientific classification Kingdom: Animalia
Tenrec
Football tournament
1951/52 Winchester City Reserves 5-2 Austin Sports The Dell 1952/53 Old Bartonians 1-0 Southampton Gas Employees The Dell 1953/54 Harland & Wolff 2–0 Winchester
Southampton_Senior_Cup
Extinct family of mammals
American Naturalist. 23: 1–29. Hooker, J. J. (1986). "Mammals from the Bartonian (middle/late Eocene) of the Hampshire Basin, southern England". Bulletin
Anoplotheriidae
Extinct family of mammals in the order Carnivora
Nimravidae is an extinct family of carnivorans, sometimes known as false saber-toothed cats, whose fossils are found in North America, Africa, and Eurasia
Nimravidae
Alienocarapus Gen. et sp. nov Valid Schwarzhans, Stringer & Takeuchi Eocene (Bartonian) Mission Valley Formation United States ( California) A member of the
2024_in_paleoichthyology
BARTONIAN
BARTONIAN
BARTONIAN
BARTONIAN
Boy/Male
Tamil
Nishikesh | நீஷீகேஷÂ
Boy/Male
German, Polish
Famous Landowner
Boy/Male
Biblical
The only Lord.
Boy/Male
Bengali, Gujarati, Hindu, Indian, Kannada, Kashmiri, Malayalam, Marathi, Sanskrit, Tamil, Telugu, Traditional
Lotus Flower
Girl/Female
Australian, Gaelic
Valley; Secluded Valley; From the Glen
Girl/Female
American, Australian, German, Scandinavian, Spanish, Swedish
Peaceful Ruler; Female Version of Eric; Ruler Forever; Rich; Strong Power; Hardy Power
Girl/Female
Muslim
Joy, Happiness, New beginning
Girl/Female
Hindu, Indian
Flower Name
Boy/Male
American, Anglo, British, English
Steward
Boy/Male
Tamil
(Father of draupad)
BARTONIAN
BARTONIAN
BARTONIAN
BARTONIAN
BARTONIAN