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Clade of animals comprising vertebrates and tunicates
vertebrate generation, gave rise to nostrils. The name Olfactores comes from Latin *olfactores ("smellers," from purposive supine olfactum of olfacio
Olfactores
Phylum of animals having a dorsal nerve cord
gills or a distinct head. The vertebrates and tunicates compose the clade Olfactores, which is sister to Cephalochordata (see diagram under Phylogeny). Extinct
Chordate
and Vertebrata are in the unranked Olfactores clade, while the subphylum Cephalochordata is not. Animals in Olfactores are characterized as having a more
List_of_chordate_orders
Subphylum of chordates
the tunicates + vertebrates form a clade, the Olfactores, with Cephalochordata as its sister (the Olfactores hypothesis), as shown in the following phylogenetic
Vertebrate
Extinct genus of tunicate
Burykhia; Therefore it is possible that it is actually a more basal group of Olfactores. The Megasiphon authors postulate the organism as a stem tunicate, rather
Megasiphon
Superphylum of bilateral animals
Deuterostomia Phylum Chordata Subphylum Cephalochordata (lancelets) Clade Olfactores Subphylum Tunicata (tunicates) Subphylum Vertebrata Infraphylum "Agnatha"
Deuterostome
Phylum of marine deuterostome animals
Deuterostomia Chordata Cephalochordata Olfactores Tunicata Vertebrata/Craniata Ambulacraria Echinodermata Hemichordata
Hemichordate
water vascular system, ambulacrum, and tube feet. However, the clade Olfactores was first proposed as part of the calcichordate theory, and has since
Calcichordate_hypothesis
Gill-bearing non-tetrapod aquatic vertebrates
fish Scientific classification Kingdom: Animalia Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Groups included "Agnatha" †"Placodermi" †"Acanthodii"
Fish
Group of vertebrates consisting of amphibians and fish
Scientific classification Kingdom: Animalia Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Groups included All non-amniote vertebrates: "Agnatha"
Anamniotes
Subphylum of chordates
Chordata Olfactores Lancelet? †Cathaymyrus? († 518 mya) †Palaeobranchiostoma? († 273.01 mya) Lancelet (crown group) Asymmetron ⊞ ⊟ Asymmetron inferum Asymmetron
Lancelet
Species of tunicate
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Thetys_vagina
Marine animals, subphylum of chordates
invertebrates. The clade consisting of tunicates and vertebrates is called Olfactores. The Tunicata contain roughly 3,051 described species, traditionally divided
Tunicate
Extinct Cambrian group of animals
showing each possible placement for Vetulicolia considered in the paper; note the presence of the Euchordata clade rather than the now-favored Olfactores.
Vetulicolia
Class of ray-finned bony fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Actinopterygii
Class of lobe-finned fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Coelacanth
Diverse clade of vertebrate animals
actinopterygians) Scientific classification Kingdom: Animalia Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Infraphylum: Gnathostomata Clade: Eugnathostomata
Osteichthyes
Infraphylum of vertebrates
Scientific classification Kingdom: Animalia Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Infraphylum: Gnathostomata Gegenbauer, 1874 Subgroups
Gnathostomata
Cartilaginous fish in the order Chimaeriformes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Chimaera
Extinct agnathan chordates resembling eels
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Conodont
Subclass of fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Elasmobranchii
Family of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Salp
Extinct family of jawless fishes
limited swimming ability. Other early vertebrates or transitional between olfactores and vertebrates are: Nuucichthys (Drumian) Emmonsaspis (Cambrian Stage
Myllokunmingiidae
Origin and diversification of fish through geologic time
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Evolution_of_fish
Extinct order of cartilaginous fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Petalodontiformes
Genus of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Pegea
Genus of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Sigillina
Extinct genus of jawless fishes
e Anaspida Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Anaspida Cowielepis Kerreralepis Pharyngolepis Pterygolepis
Pharyngolepis
Extinct genus with possible tunicate affinities
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Cheungkongella
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Synoicum
Paraphyletic infraphylum of fish
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Agnatha
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Eudistoma
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Kukenthalia
Extinct genus of jawless fishes
e Anaspida Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Anaspida Cowielepis Kerreralepis Pharyngolepis Pterygolepis
Cowielepis
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Diplosoma_(tunicate)
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Placentela
Clade of lobe-finned fish
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Sarcopterygii
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Podostyela
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Hypobythius
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Clavelina
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Oligotrema
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Morchellium
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Culeolus
Family of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Perophoridae
Genus of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Cystodytes
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Bostrichobranchus
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Pareugyrioides
Cambrian fossil chordate
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Yunnanozoon
Extinct genus of jawless fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Haikouichthys
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Perophora
Extinct genus of jawless vertebrates
e Anaspida Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Anaspida Cowielepis Kerreralepis Pharyngolepis Pterygolepis
Jamoytius
Division or infraclass of fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Teleost
Chordata Chordates (Vertebrates and closely related invertebrates) 530 Mya Olfactores Excludes cephalochordates (Lancelets) Subphylum Vertebrata Fish / Vertebrates
Timeline_of_human_evolution
Family of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Diazonidae
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Lists_of_prehistoric_fish
Class of jawed cartilaginous fish
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Chondrichthyes
Extinct family of possible tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Ausiidae
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
List of prehistoric cartilaginous fish genera
List_of_prehistoric_cartilaginous_fish_genera
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Phallusia
Family of fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Snailfish
Extinct group of elasmobranchs
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Phoebodontiformes
Clade of ray-finned fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Cladistia
Family of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Octacnemidae
Larvacean genus
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Fritillaria_(tunicate)
Extinct genus of jawless fishes
e Anaspida Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Anaspida Cowielepis Kerreralepis Pharyngolepis Pterygolepis
Birkenia
Extinct family of jawless fishes
e Anaspida Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Anaspida Cowielepis Kerreralepis Pharyngolepis Pterygolepis
Euphaneropidae
Order of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Doliolida
Paraphyletic class of tunicates comprising sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Ascidiacea
Clade of vertebrates
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Rhipidistia
Extinct paraphyletic class of fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Placoderm
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Megalodicopia
Extinct genus of primitive chordates
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Pikaia
Extinct paraphyletic class of fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Acanthodii
Subclass of fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Chondrostei
Species of Appendicularia
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Bathochordaeus_mcnutti
Family of jawless fishes
Osteostraci Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Osteostraci Hemicyclaspis Ungulaspis Ateleaspididae
Ateleaspididae
Group of extinct jawless vertebrates
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Anaspida
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Claudenus
Extinct class of jawless fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Pteraspidomorpha
Extinct group of cartilaginous fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Eugeneodontiformes
Larvacean genus
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Kowalevskia
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Sidneioides
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Botryllocarpa
Genus of larvacean
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Mesochordaeus
Superclass of jawless fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Cyclostomi
Genus of jawless fishes
e Anaspida Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Anaspida Cowielepis Kerreralepis Pharyngolepis Pterygolepis
Lasanius
Extinct order of cartilaginous fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Ctenacanthiformes
Family of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Polyclinidae
Genus of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Polyzoa_(tunicate)
Genus of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Polysyncraton
Family of sea squirts
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Ascidiidae
Extinct order of cartilaginous fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Xenacanthiformes
Extinct genus of chordates
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Cathaymyrus
Extinct genus of jawless fishes
e Anaspida Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Anaspida Cowielepis Kerreralepis Pharyngolepis Pterygolepis
Cornovichthys
Class of tunicates
Tunicate genera Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Tunicata Appendicularia (Larvacea) Kowalevskia Fritillariidae Appendicularia
Thaliacea
Armored jawless fish of the Paleozoic
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Ostracoderm
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
List_of_placoderm_genera
Genus of prehistoric jawless fish
Osteostraci Kingdom: Animalia Clade: Bilateria Phylum: Chordata Clade: Olfactores Subphylum: Vertebrata Osteostraci Hemicyclaspis Ungulaspis Ateleaspididae
Tremataspis
Subclass of fishes
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Neopterygii
Extinct genus of chordates
Forerunners Basal member †Pikaia Cephalochordate †Cathaymyrus Lancelet Olfactores †Haikouella Tunicate †Myllokunmingiidae? (†Haikouichthys, †Myllokunmingia)
Zhongxiniscus
OLFACTORES
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