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Example of an organism used to describe its species
A holotype is a single physical example (or illustration) of an organism that is the one that was used when the species (or lower-ranked taxon) was formally
Holotype
Carcharodontosaurid dinosaur genus from the late Cretaceous period
Late Cretaceous period, approximately 99.6 to 95 million years ago. The holotype specimen was discovered in the Candeleros Formation of Patagonia in 1993
Giganotosaurus
Extinct genus of dinosaurs
Oviraptor is only known from a single partial skeleton regarded as the holotype, as well as a nest of about fifteen eggs and several small fragments from
Oviraptor
Extinct genus of carnivorous ungulate from Eocene epoch
fossa suggest that Andrewsarchus likely had a fairly weak bite force. The holotype of Andrewsarchus mongoliensis is a mostly complete cranium (specimen number
Andrewsarchus
Spinosaurid theropod dinosaur genus from the Early Cretaceous Period
until its return to Brazil in 2026. In 1996, the specimen became the holotype of the type species Irritator challengeri. The genus name comes from the
Irritator
Genus of theropod dinosaurs
Surrey, England, in sediments of the Weald Clay Formation, and became the holotype specimen of Baryonyx walkeri, named by palaeontologists Alan J. Charig
Baryonyx
subrectangular in outline. The holotype was once tentatively assigned to "Coelosaurus" antiquus. Shark bites present on the holotype suggest that the specimen's
List of informally named dinosaurs
List_of_informally_named_dinosaurs
Ankylosaurid dinosaur genus from the Late Cretaceous Period
teeth of two skulls assigned to A. magniventris differed from those of the holotype specimen in some details, and though he expressed a "considerate temptation"
Ankylosaurus
Specimen(s) chosen to represent a newly described taxon
nomenclatural type when there was no holotype specified or the holotype has been lost or destroyed isotype – a duplicate of the holotype syntype – any specimen (or
Type_(biology)
Genus of theropod dinosaur from the Late Jurassic period
Marsh, but more likely was used solely for display. The first specimen, holotype USNM 4735, was discovered and excavated by farmer Marshall Parker Felch
Ceratosaurus
Extinct genus of sperm whale from the Miocene epoch
suitable for hunting large prey. The body length of Livyatan melvillei holotype is approximately 13.5–17.5 meters (44–57 ft) long. Comparable to that of
Livyatan
Genus of theropod dinosaurs
of Nanotyrannus, as histology of the hyoids indicates the N. lancensis holotype belongs to a mature individual. Nanotyrannus was a mid-sized tyrannosauroid
Nanotyrannus
Extinct genus of birds
Aguirre-Zabala in Comallo, in the region of Patagonia, and was made the holotype of the new genus and species Kelenken guillermoi in 2007. The genus name
Kelenken
Extinct genus of ants
slightly smaller in size then the coeval species. The Titanomyrma lubei holotype was discovered by Louis Lube at DMNS locality 784, the Farson Fish Beds
Titanomyrma
Extinct genus of reptiles
briefly four years earlier in a paper on Lufeng fossil vertebrates. The holotype jaw consisted of 11 alveoli (tooth sockets) on the left side and 12 alveoli
Platyognathus
Genus of moths
Stegothyris fasciculalis, which is found in Madagascar and South Africa. The holotype of 'Stenia fasciculalis' Zeller, 1852 is illustrated on the web pages of
Stegothyris
Genus of pteranodontoid pterosaur from the Early Cretaceous
Yixian Formation of Liaoning, China. It is known from a single specimen, holotype JZMP-V-12, housed at the Benxi Geological Museum. This specimen was modified
Yixianopterus
Extinct genus of marine reptiles
specimen. Based on several distinctive features, he decided to make it the holotype of a subgenus accompanying a new species of Plesiosaurus, which he named
Pliosaurus
Extinct genus of dinosaurs
indicates a generalist diet with a potentially occasional carnivory. The holotype—and only known specimen—has been determined to represent a young adult
Gigantoraptor
Extinct genus of mammals
Auroracetus, from Aurora (the location the holotype was discovered, Aurora N.C.) and the Latin cetus (whale), is a genus of Pontoporiidae, the river dolphins
Auroracetus
Specimen that helps define what a taxon represents
name of a species and other taxon actually represents, but it is not the holotype (and in botany is also neither an isotype nor a syntype). Often there is
Paratype
Extinct chimaeric genus of dinosaurs
D. steini were discovered in a multi-species bonebed. Elements of the holotype and referred specimens were later found to belong to trionychid turtles
Dakotaraptor
Extinct genus of therapsids
related to mammals than to dinosaurs, the latter being sauropsids. The holotype and only known specimen of Dinosaurus was discovered in the early 1840s
Dinosaurus
Genus of crayfishes
Palaeoechinastacus is known from one partial body fossil, which is the holotype, and has been deposited in the National Museum of Victoria in Melbourne
Palaeoechinastacus
Sauropod dinosaur genus from the late Jurassic Period
and 22 meters (59 and 72 ft) long; body mass estimates of the subadult holotype specimen range from 28.3 to 46.9 metric tons (31.2 to 51.7 short tons)
Brachiosaurus
Genus of theropod dinosaur from the Early Jurassic
preserved were collected in 1942. The most complete specimen became the holotype of a new species in the genus Megalosaurus, named M. wetherilli by Samuel
Dilophosaurus
Extinct genus of dinosaurs
University of Kansas, Yale University, and the University of New Orleans. The holotype fossil is less than 1 m (3.3 ft) long, although this specimen appears to
Bambiraptor
Extinct genus of dinosaurs
that a more thorough review of the specimens is required. Because the holotype of T. formosus is a single tooth, this renders Troodon a nomen dubium.
Troodon
Genus of abelisaurid dinosaurs
hard, in reference to the heavily cemented rocks in which the holotype was found. The holotype consists of three caudal vertebrae and a partial pelvic girdle
Kurupi_itaata
Sauropod dinosaur genus from Late Jurassic-Early Cretaceous Period
mounting, and description of the skeleton were completed in 1965. The holotype specimen (CCG V 20401) consisted of an almost complete and articulated
Mamenchisaurus
Genus of theropod dinosaurs
in the Nemegt Formation of Mongolia. In 1970, this specimen became the holotype of the only species within the genus, Deinocheirus mirificus; the genus
Deinocheirus
tenuis, Davis 1887 Type Locality: Sendenhorst (Westphalia, Germany) The Holotype is specimen "GIM-8434". As reported in Dietze's study : "Complete specimen
Telepholis
Species of damselfly
refers to the fact that the species existed in the Cretaceous period. The holotype, NHMUK CH 879vii, a fossilized wing, was found n lagoonal sediments in
Steleopteron_cretacicus
Extinct genus of jawless fishes
among the earliest known vertebrate animals in the fossil record. The holotype of the type species, Myllokunmingia fengjiaoa — and the only uncontroversial
Myllokunmingia
Extinct genus of mammals
plate. The first upper incisor presents a groove in its lingual part. The holotype, SGOPV 5157, held by the Chilean National Museum of Natural History in
Eotypotherium
Extinct subspecies of carnivoran
lower carnassial (m1) tooth excavated from deposits in Kuruwita as the holotype. It was recorded as specimen BMNH M51883 in the British Museum Natural
Panthera_leo_sinhaleyus
Extinct genus of dragonfly-like insects
a body length from head to tail of almost 430 millimetres (17 in). The holotype is held in the Museum of Comparative Zoology. Meganeuropsis americana,
Meganeuropsis
Extinct genus of birds
occurrence of phorusrhacids during the Pleistocene epoch. The Eschatornis holotype specimen, MCL-A-1.795, was discovered and collected by Cástor Cartelle
Eschatornis
Pliosaur genus from the Early Cretaceous period of Australia
1924 that Albert Heber Longman formally described this specimen as the holotype of an imposing pliosaurid, to which he gave the scientific name K. queenslandicus
Kronosaurus
Extinct genus of reptiles
carolinensis, meaning "Carolina butcher". Two specimens are known, the holotype skull and skeleton NCSM 21558, and the referred humerus NCSM 21623. The
Carnufex
Extinct species of shrew
on the paleontology of mammals in the southwestern United States. The holotype specimen is deposited at the University of Texas at El Paso Biodiversity
Notiosorex_harrisi
Extinct genus of theropod dinosaurs
least two unrelated taxa. Some of the Eocarcharia material, including the holotype (name-bearing) specimen, likely belongs to a baryonychine spinosaurid.
Eocarcharia
Genus of extinct mammals
femur). The holotype was found in a silt and mudstone bed over a 1.8 m2 (19 sq ft) area. Further excavation recovered most of the holotype's skeleton—most
Ambulocetus
Species of shrub endemic to Australia
area in New South Wales, Australia. It is currently only known from the holotype specimen. Grevillea milleriana is a low, spreading, sub-prostrate shrub
Grevillea_milleriana
Taxonomy of a dinosaur genus
coined the name Allosaurus fragilis in 1877 for a fragmentary specimen (the holotype specimen; the specimen on which the species was based). According to Marsh
Taxonomy_of_Allosaurus
Extinct genus of dinosaurs
where it was brought for preparation. In 2004, the specimen became the holotype of the new genus and species Atrociraptor marshalli; the generic name is
Atrociraptor
Genus of Cretaceous cephalopods
traditionally recognized. In 2019, due to the poor preservation of the holotype specimen, Tusoteuthis was theorized to likely be a nomen dubium, and it
Tusoteuthis
Extinct genus of dinosaur
the fossils and enhanced virtual and long-distance collaboration. The holotype specimen, MPM-PV 1156, consists of a partial skeleton, somewhat preserved
Dreadnoughtus
Extinct genus of reptiles
exhumation. In 1903, Samuel Wendell Williston designated the fossil as the holotype of a new plesiosaur genus and species under the name Brachauchenius lucasi
Brachauchenius
Extinct genus of dinosaurs
Provence at the foot of which the animal's remains were unearthed. The holotype specimen, MNHN BO001, consists of the second toe claw of the left foot
Pyroraptor
Extinct genus of dinosaurs
sauropod dinosaur from the Late Cretaceous (Maastrichtian) of Romania. The holotype of this genus was originally referred to Magyarosaurus hungaricus, which
Uriash
Extinct genus of dinosaurs
paleontological collections of the Musée National Boubou Hama in Niamey. The holotype, catalogued as MNN GDF500, consists of a partial skeleton lacking the skull
Suchomimus
Extinct genus of dinosaurs
it was a herrerasaurid or herrerasaurian. A 2019 redescription of its holotype considered Chindesaurus to be a theropod closely related to Tawa, a slightly
Chindesaurus
Species of gastropod
Costellariidae, the ribbed miters. The length of the shell attains 43.7 mm. The holotype of the species (MNHN IM-2000-30178), measured 42.2 mm. Type Locality :
Tosapusia_sauternesensis
Species of mollusc
from the Americas (North America and South America). The holotype collected was named holotype MZSP 155717 and the paratype MZSP 155716. All were discovered
Eupera_troglobia
Extinct genus of reptiles
(Barremian) Yixian Formation in Liaoning, China. A redescription of the holotype proposed affinities of Liaoningotitan with the Euhelopodidae. Liaoningotitan
Liaoningotitan
Extinct species of snake
extinct python species. It lived during the early/middle Miocene. The holotype is a single trunk vertebra found in France. Fossilized remains of Python
Python_europaeus
Tyrannosaurid theropod dinosaur genus from Late Cretaceous period
alongside the Sinankylosaurus holotype in 2008, and is currently housed at the Zhucheng Dinosaur Museum. Casts of the holotype, IVPP FV 1794, are held at
Zhuchengtyrannus
Extinct species of carnivore
Pleistocene. It is named after the Gamssulzen Cave in Austria, where the holotype of this species was found. Ursus ingressus was a large cave bear with massive
Ursus_ingressus
Extinct genus of mammals
in the upper molars and of the trigonid fossa in the lower molars. The holotype MLP 88-V-10-6 is a fragmentary mandible preserving the second molar, kept
Punahyrax
Extinct genus of reptiles
similarities to the holotype of Qiupalong. The foot claws of the new specimens in particular were nearly identical to those of the holotype. As a result, they
Qiupalong
Genus of ankylosaurid dinosaur
Granger, who accompanied the 1923 expedition as a paleontologist. The holotype, AMNH 6523, was found in a layer of the Djadokhta Formation, dating from
Pinacosaurus
Species of moth
a moth of the family Cossidae. It is found in eastern Madagascar. The holotype had been found in Ifanadiana in 1995. List of moths of Madagascar Yakovlev
Planctogystia_lemur
Extinct genus of turtles
thalassochelydian turtle. Its type species is Eurysternum wagleri, the holotype of which has since been lost and only survives in illustrations. Eurysternum
Eurysternum
Genus of reptiles (fossil)
Canada. Gryphoceratops is known only from the holotype ROM 56635, a partial right dentary. The holotype was collected in the northwest corner of Dinosaur
Gryphoceratops
Genus of sauropod dinosaurs
titanosauriforms. The holotype of Ninjatitan was found at the same locality, but a slightly deeper rock layer, as the holotype of the diplodocoids Leinkupal
Ninjatitan
Extinct genus of dinosaurs
many complete specimens, including the nearly complete holotype found by Brown in 1911. The holotype skeleton is only missing the last section of the tail
Corythosaurus
Extinct species of kiwi
Zealand's North Island. The species was first described in 2021 based on the holotype (NMNZ S.36731), a complete left tarsometatarsus that was found in the Pleistocene
Apteryx_littoralis
Genus of algae
the species Pycnococcus provasolii. This spieces is the type species (holotype) of the genus Pycnococcus. Guiry, M.D.; Guiry, G.M. "Pycnococcus". AlgaeBase
Pycnococcus
Genus of frond-like lifeforms
the discovery in the Journal of the Yorkshire Geological Society. The holotype (the actual physical example from which the species was first described)
Charnia
Extinct species of reptile
and dinosaurs. Kadimakara is only known from parts of the skull. The holotype specimen, QMF 6710, includes the rear part of the skull and a fragment
Kadimakara_australiensis
Species of beetle
species of carabid beetle named after the actress Catherine Bell. The holotype was collected in Costa Rica and first described to science in 2002. Terry
Agra_catbellae
Species of beetle
Cerambycidae. It was described by Francis Polkinghorne Pascoe in 1865. The holotype was collected by Alfred Russel Wallace in Waigeo. Pascoe, Francis P. (1865)
Ropica_tentata
Extinct genus of dinosaurs
ornamentation along the edge of this embayment. The specimen that would become the holotype (name-bearing specimen) of Cryptarcus was discovered in 1936 by palaeontologist
Cryptarcus
Species of fly
black. The legs are black, except at the tibiae, which are yellow. The holotype, named as Villa webberi by Johnson, 1919, is a male found at Massachusetts
Hemipenthes_webberi
Genus of theropod dinosaurs
approaching a similar size. The type specimen of Austroraptor cabazai, holotype MML-195, was recovered in the Bajo de Santa Rosa locality of the Allen
Austroraptor
Species of bat
is a species of vesper bat in the genus Murina. It is only known by the holotype, an old female, collected in an abandoned mine on Tsushima Island in 1962
Gloomy_tube-nosed_bat
Genus of azhdarchid pterosaur
two specimens, the holotype, consisting of a partial postcranial skeleton, and the paratype, consisting of a left humerus. The holotype includes material
Thanatosdrakon
Extinct genus of dinosaurs
Rhaetian) in what is now Telangana, central India. It is known from the holotype ISI R273, parts 1–3, partially articulated postcranial material and from
Nambalia
Species of moth
Per Olof Christopher Aurivillius in 1910. It is found in Tanzania. The holotype of this species is illustrated on the web-pages of the Swedish Museum of
Parotis_fasciculata
Extinct genus of eurypterid
and Latin arachne meaning "spider", translates to "great spider". The holotype (now stored at the Museum of Paleontology at the National University of
Megarachne
Extinct genus of reptiles
differences with the holotype specimen. Noting this, McGowan redescribed this specimen in more detail and made it the holotype of an entirely new species
Temnodontosaurus
Extinct population of the brown bear
Southwestern United States and Mexico. The specimen later designated the holotype of U. a. nelsoni was shot by H. A. Cluff at Colonia Garcia, Chihuahua,
Mexican_grizzly_bear
Place where a name-bearing type specimen was collected
define "type locality" as a regulated term, the collecting locality of the holotype or other original material is routinely referred to by botanists as the
Type_locality_(biology)
Genus of snakes
at the Museum of Zoology (UNAM) as MZFC-HE 37100, and assigned as the holotype of Y. tlalli. It measures 38.1 centimetres (15.0 in) in total length. A
Yakacoatl
Species of cartilaginous fish
however the holotype specimen of B. leucomelanos was caught on the Coriolis Bank, off western New Caledonia in 2002. The male holotype specimen measures
Bathyraja_leucomelanos
Species of lizard
Agamidae. The species is endemic to Indonesia. Once only known from the holotype, a single specimen collected from North Sumatra in 1891, the species was
Harpesaurus_modiglianii
Extinct genus of dinosaurs
Archaeopteryx. Three specimens have been assigned to Sinosauropteryx prima: the holotype GMV 2123 (and its counter slab [opposite face], NIGP 127586), NIGP 127587
Sinosauropteryx
Extinct genus of dinosaurs
discovered material, referred to as DMNH 21461, is now recognized as the holotype of Nanuqsaurus. As well as this, the initial discoveries of teeth were
Nanuqsaurus
Extinct species of reptile
be clenched between the teeth of BMNH R1157 (its holotype) may in fact belong to a victim. The holotype, obtained from the Lower Lias of Lyme Regis, was
Temnodontosaurus_eurycephalus
Extinct genus of dinosaurs
Cretaceous) of Argentina. The type and only species is N. leali. The fragmentary holotype specimen of Noasaurus, PVL 4061, consisting of a few cranial and postcranial
Noasaurus
Extinct species of bird
published in 2001. The holotype is in the collection of the Museum of New Zealand Te Papa Tongarewa. It is known from the holotype tarsometatarsus and paratypes
Viti_Levu_giant_pigeon
Extinct Pliosaurid marine reptile
mainly known from Europe, with one occurrence reported in Mexico. As the holotype specimen of L. ferox consists of a single tooth preserving questionable
Liopleurodon
Extinct genus of dinosaurs
midline of the skull outwards to the top end of the orbit. The Gremlin holotype specimen, TMP 2011.053.0027, was discovered in sediments of the Oldman
Gremlin_slobodorum
Species of gastropod
gastropod mollusk, in the family Costellariidae, the ribbed miters. The holotype of the species (MNHN IM-2013-19695), measured 35.1 mm. Type Locality :
Tosapusia_bismarckeana
Species of moth
Hungarian lepidopterologist who collected the holotype. The wingspan is about 22 millimetres (0.87 in) (holotype, a female). Mironov, V. & Galsworthy, A. C
Eupithecia_szaboi
Extinct genus of dinosaurs
contains only one known species, Halszkaraptor escuilliei. The type specimen (holotype) has been compared to the bones of extant crocodilians and aquatic birds
Halszkaraptor
Extinct species of mammal
described by the Dutch paleoanthropologist Eugène Dubois in 1908. The holotype of Bos palaesondaicus is a skull from Trinil. This species is the likely
Bos_palaesondaicus
Extinct genus of dinosaurs
were. Sinornithosaurus was among the smallest dromaeosaurids, with the holotype measuring 1.2 metres (3.9 ft) long and weighing 3–5 kilograms (6.6–11.0 lb)
Sinornithosaurus
Species of gastropod
terrestrial pulmonate gastropod mollusk in the family Polygyridae. The holotype was collected on May 24, 1975 from talus accumulation of igneous rock on
Ashmunella_todseni
HOLOTYPE
HOLOTYPE
HOLOTYPE
HOLOTYPE
Boy/Male
Indian, Punjabi, Sikh
Victory of Beauty
Girl/Female
Indian
Boy/Male
Tamil
Alolupan | அலோலà¯à®ªà®£
One of the kauravas
Boy/Male
Indian
Protractor, One who worships God
Girl/Female
Indian, Tamil
Beautiful Like a Pearl
Girl/Female
Arabic, Muslim
Light of the Day Break; The Light at Dawn; Morning Light
Boy/Male
British, English
Son of the Highborn
Surname or Lastname
English
English : variant of Bolding.Swedish (Boldén) : ornamental name.
Girl/Female
Tamil
Queen of music and songs
Boy/Male
Gujarati, Hindu, Indian, Punjabi, Sikh, Tamil
Knowledgeable
HOLOTYPE
HOLOTYPE
HOLOTYPE
HOLOTYPE
HOLOTYPE